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Aminooxyacetic acid inhibits the malate-aspartate shuttle in isolated nerve terminals and prevents the mitochondria from utilizing glycolytic substrates

In drugs that have particularly dangerous side effects, the FDA can require a boxed warning, otherwise known as a black box warning

Since superoxide anion cannot diffuse across the inner mitochondrial membrane it remains in the matrix which allows it to react with Fe-S groups in mitochondrial proteins resulting in the liberation of ferrous (Fe 2+ ) iron

However, studies using gas chromatography/mass spectrometry (GC/MS) used the same GC-like properties